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{{Taxobox
| name = Burung hantu bertandukBeluk
| image = Bengalese Eagle Owl.jpg
| image_caption = ''Bubo bengalensis'' Burung hantu bertanduk dari Indian
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''Ophthalmomegas'' <small>Dejaut, 1911</small><ref>Possibly a [[junior synonym]] of ''Ketupa'', if that is a valid genus: Pavia (1999), Mlíkovský (2002, 2003).</ref>
}}
'''Burung hantu bertandukBeluk''' Amerika ;merupakan jenis burung hantu bertanduk dari [[genus]] '''''Bubo'''''. Genus ini memiliki sekitar satu atau dua lusin [[species]] dari keluarga [[burung hantu]], dan dapat ditemukan di berbagai tempat di dunia. Pada awalnya hanya burung hantu yang memiliki jumbai pada telinganya yang dimasukkan ke genus ini.
[[Berkas:White horned owl portrait.jpg|thumbjmpl|Seekor burung hantu bertanduk putih sedang menyantap seekor tikus.]]
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==Systematics==
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[[mtDNA]] [[cytochrome b|cytochrome ''b'']] [[DNA sequence|sequence]] data supports the decision to consider the [[Snowy Owl]] an eagle-owl [[adaptation|adapted]] to [[Arctic]] conditions and moving it into ''Bubo'', rendering the [[monotypic]] genus ''Nyctea'' invalid.<ref name = olsenetal2002>Olsen ''et al.'' (2002)</ref>
 
The four fish-owls previously in the genus ''[[Ketupa]]'' were provisionally moved into ''Bubo'' as well.<ref>König ''et al.'' (1999)</ref> However, the mtDNA cytochrome ''b'' data suggests that in this case, to make ''Bubo'' [[monophyletic]] the ''[[Scotopelia]]'' fishing owls would also need to be included there. On the other hand, the genus then becomes quite large and ill-defined, and ''Bubo'' in the expanded sense seems to consist of two distinct [[clade]]s. Thus, the fish and fishing owls can alternatively be united in ''Ketupa'' if some aberrant eagle-owls &ndash; at least the [[Barred Eagle-Owl|Barred]], [[Spot-bellied Eagle-Owl|Spot-bellied]] and [[Usambara Eagle-Owl]]s, perhaps also [[Fraser's Eagle-Owl]] and maybe others &ndash; are moved into that genus too. As some enigmatic eagle-owls remain essentially unstudied and others &ndash; e.g. [[Verreaux's Eagle-Owl]] &ndash; are of unresolved relationships, more research is needed.<ref name = olsenetal2002 />
 
===Living species===
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* ''Bubo leakeyae'' (Early Pleistocene of Tanzania)
* ''Bubo binagadensis'' (Late Pleistocene of Binagady, Azerbaijan)
* [[Cuban Horned Owl]], ''[[Bubo osvaldoi]]'' (Late{{Verify source|date=September 2008}} Pleistocene of Cuba)
 
Some notable undescribed [[fossil]]s of prehistoric horned owls, usually quite fragmentary remains, have also been recorded:
Baris 79:
Several presumed ''Bubo'' fossils have turned out to be from different birds. The Late Eocene/Early Oligocene eared owls ''"Bubo" incertus'' and ''"Bubo" arvernensis'' are now placed in the fossil [[barn-owl]] genera ''[[Nocturnavis]]'' and ''[[Necrobyas]]'', respectively. ''"Bubo" leptosteus'' is now recognized as primitive owl in the genus ''[[Minerva (owl)|Minerva]]'' (formerly ''Protostrix''). ''"Bubo" poirreiri'' from the Late Oligocene or Early Miocene of Saint-Gérard-le-Puy in France, is now placed in ''[[Mioglaux]]''.
 
On the other hand, the supposed fossil [[heron]] ''"Ardea" lignitum'' from the [[Late Pliocene]] of [[Plaue]]-Rippersroda (Germany) was apparently an owl and close to ''Bubo'' or more probably actually belongs here. Given its age &ndash; about 2 million years ago or so &ndash; it is usually included in the Eurasian Eagle-Owl today.<ref>Olson (1985): p.167, Mlíkovský (2002)</ref>
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== Catatan kaki ==
{{Reflist|colwidth=30em}}
 
== Referensi ==
* [[Alan Feduccia|Feduccia, J. Alan]] (1970): Some birds of prey from the Upper Pliocene of Kansas. ''[[Auk (journal)|Auk]]'' '''87'''(4): 795-797. [http://elibrary.unm.edu/sora/Auk/v087n04/DJVU/P0795-P0797.djvu DjVu fulltext] {{Webarchive|url=https://web.archive.org/web/20110606171151/http://elibrary.unm.edu/sora/Auk/v087n04/DJVU/P0795-P0797.djvu |date=2011-06-06 }} [http://elibrary.unm.edu/sora/Auk/v087n04/p0795-p0797.pdf PDF fulltext] {{Webarchive|url=https://web.archive.org/web/20110606111215/http://elibrary.unm.edu/sora/Auk/v087n04/p0795-p0797.pdf |date=2011-06-06 }}
* Howard, Hildegarde (1947): A preliminary survey of trends in avian evolution from Pleistocene to recent time. ''[[Condor (journal)|Condor]]'' '''49'''(1): 10–13. [http://elibrary.unm.edu/sora/Condor/files/DJVU/v049n01/P0010-P0013.djvu DjVu fulltext] {{Webarchive|url=https://web.archive.org/web/20130702224747/http://elibrary.unm.edu/sora/Condor/files/DJVU/v049n01/P0010-P0013.djvu |date=2013-07-02 }} [http://elibrary.unm.edu/sora/Condor/files/issues/v049n01/p0010-p0013.pdf PDF fulltext] {{Webarchive|url=https://web.archive.org/web/20081217155256/http://elibrary.unm.edu/sora/Condor/files/issues/v049n01/p0010-p0013.pdf |date=2008-12-17 }}
* König, Claus; Weick, Friedhelm & Becking, Jan-Hendrik (1999): ''Owls: A guide to the owls of the world.'' [[Yale University Press]], New Haven. <small>ISBN 0-300-07920-6</small>
* [[Kalman Lambrecht|Lambrecht, Kálmán]] (1933): ''Handbuch der Palaeornithologie'' [Handbook of Paleornithology]. Gebrüder Bornträger, Berlin. [in German]
* Mlíkovský, Jiří (2002): ''Cenozoic Birds of the World, Part 1: Europe''. Ninox Press, Prague. <small>{{Listed Invalid ISBN|80-901105-3-8}}</small> [http://www.nm.cz/download/JML-18-2002-CBE.pdf PDF fulltext] {{Webarchive|url=https://web.archive.org/web/20110520101755/http://www.nm.cz/download/JML-18-2002-CBE.pdf |date=2011-05-20 }} {{Webarchive|url=https://www.webcitation.org/5wzfLX4u8?url=http://www.nm.cz/download/JML-18-2002-CBE.pdf |date=2011-03-07 }}<!-- This should be treated with extreme caution as regards merging of species. Splits are usually good though. See also critical review in Auk121:623-627 here http://www.findarticles.com/p/articles/mi_qa3793/is_200404/ai_n9396879 -->
* Mlíkovský, Jiří (2003): Brown Fish Owl (''Bubo zeylonensis'') in Europe: past distribution and taxonomic status. ''Buteo'' '''13''': 61-65. [http://www.nm.cz/download/JML-13-2003-bubo-zeylonensis.pdf PDF fulltext] {{Webarchive|url=https://web.archive.org/web/20070927223329/http://www.nm.cz/download/JML-13-2003-bubo-zeylonensis.pdf |date=2007-09-27 }}
* Olsen, Jery; Wink, Michael; Sauer-Gürth, Heidi & Trost, Susan (2002): A new ''Ninox'' owl from Sumba, Indonesia. ''[[Emu (journal)|Emu]]'' '''102'''(3): 223-231. <small>{{DOI|10.1071/MU02006}}</small> [http://www.uni-heidelberg.de/institute/fak14/ipmb/phazb/pdf-files/2002%20Pdf.Pubwink/17.2002.pdf PDF fulltext] {{Webarchive|url=https://web.archive.org/web/20090930232448/http://www.uni-heidelberg.de/institute/fak14/ipmb/phazb/pdf-files/2002%20Pdf.Pubwink/17.2002.pdf |date=2009-09-30 }}
* [[Storrs Olson|Olson, Storrs L.]] (1985): The fossil record of birds. ''In:'' Farner, D.S.; King, J.R. & Parkes, Kenneth C. (eds.): ''Avian Biology'' '''8''': 79-238. Academic Press, New York.
* Pavia, Marco (1999): Un cranio di ''Bubo insularis'' Mourer-Chauviré & Weesie, 1986 (Aves, Strigidae) nelle brecce ossifere del Pleistocene di Capo Figari (Sardegna, [[Italia]]) ["A cranium of ''B. insularis'' from the Pleistocene ossiferous breccia of Cape Figari (Sardinia, Italy)"]. ''Atti della Accademia delle Scienze di Torino, Classe di Scienze fisiche, matematiche e naturali'' '''133''': 1-10 [Italian with English abstract]. [http://www2.nrm.se/ve/birds/sape/GlobalOwlProject/Fossil_owls/Pavia%201999.pdf PDF fulltext] {{Webarchive|url=https://web.archive.org/web/20120925034803/http://www2.nrm.se/ve/birds/sape/GlobalOwlProject/Fossil_owls/Pavia%201999.pdf |date=2012-09-25 }}
* [[David Steadman|Steadman, David William]]; Arroyo-Cabrales, Joaquin; Johnson, Eileen & Guzman, A. Fabiola (1994): New Information on the Late Pleistocene Birds from San Josecito Cave, Nuevo León, Mexico. ''[[Condor (journal)|Condor]]'' '''96'''(3): 577-589. [http://elibrary.unm.edu/sora/Condor/files/DJVU/v096n03/P0577-P0589.djvu DjVu fulltext] {{Webarchive|url=https://web.archive.org/web/20130702152740/http://elibrary.unm.edu/sora/Condor/files/DJVU/v096n03/P0577-P0589.djvu |date=2013-07-02 }} [http://elibrary.unm.edu/sora/Condor/files/issues/v096n03/p0577-p0589.pdf PDF fulltext] {{Webarchive|url=https://web.archive.org/web/20060911032144/http://elibrary.unm.edu/sora/Condor/files/issues/v096n03/p0577-p0589.pdf |date=2006-09-11 }}
{{Taxonbar|from=Q214293}}
 
[[Kategori:FaunaStrigiformes]]
[[Kategori:Burung|Hantu bertanduk]]