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}}|style1=font-size:80%; line-height:80%|label1=[[Hominoidea]] (hominoid, kera besar)}}
[[Berkas:Lucy_Skeleton.jpg|kiri|jmpl|Rekonstruksi terhadap [[Lucy (Australopithecus)|Lucy]]'','' kerangka ''[[Australopithecus afarensis]]'' pertama yang ditemukan.]]
Genus ''Homo'' berevolusi dari ''[[Australopithecus]]''.<ref>{{cite journal|date=September 2010|title=The Evolutionary History of the Australopiths|url=https://evolution-outreach.biomedcentral.com/articles/10.1007/s12052-010-0249-6|journal=Evolution: Education and Outreach|language=en|volume=3|issue=3|pages=341–352|doi=10.1007/s12052-010-0249-6|issn=1936-6434|archive-url=https://web.archive.org/web/20220531204050/https://evolution-outreach.biomedcentral.com/articles/10.1007/s12052-010-0249-6|archive-date=31 May 2022|access-date=30 July 2022|vauthors=Strait DS|s2cid=31979188|url-status=live}}</ref><ref>{{cite journal|date=September 2010|title=Origin of the Genus Homo|url=https://evolution-outreach.biomedcentral.com/articles/10.1007/s12052-010-0247-8|journal=Evolution: Education and Outreach|language=en|volume=3|issue=3|pages=353–366|doi=10.1007/s12052-010-0247-8|issn=1936-6434|archive-url=https://web.archive.org/web/20220523050858/https://evolution-outreach.biomedcentral.com/articles/10.1007/s12052-010-0247-8|archive-date=23 May 2022|access-date=30 July 2022|vauthors=Dunsworth HM|s2cid=43116946|url-status=live}}</ref> Meskipun [[Fosil|fosil-fosil]] dari masa transisi ini langka, anggota ''Homo'' yang paling awal memiliki beberapa ciri-ciri utama yang sama dengan ''Australopithecus''.<ref>{{cite journal|date=July 2016|title=From Australopithecus to Homo: the transition that wasn't|journal=Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences|volume=371|issue=1698|page=20150248|doi=10.1098/rstb.2015.0248|pmc=4920303|pmid=27298460|vauthors=Kimbel WH, Villmoare B|s2cid=20267830}}</ref><ref name="Villmoare2015">{{cite journal|display-authors=6|date=March 2015|title=Paleoanthropology. Early Homo at 2.8 Ma from Ledi-Geraru, Afar, Ethiopia|journal=[[Science (journaljurnal)|Science]]|volume=347|issue=6228|pages=1352–1355|bibcode=2015Sci...347.1352V|doi=10.1126/science.aaa1343|pmid=25739410|vauthors=Villmoare B, Kimbel WH, Seyoum C, Campisano CJ, DiMaggio EN, Rowan J, Braun DR, Arrowsmith JR, Reed KE|doi-access=free}}</ref> Catatan paling awal dari ''Homo'' adalah spesimen berusia 2,8 juta tahun, LD 350-1 dari Ethiopia, dan spesies yang paling awal adalah ''[[Homo habilis]]'' dan ''[[Homo rudolfensis]]'' yang berevolusi 2,3 juta tahun yang lalu.<ref name="Villmoare2015" /> ''[[Homo erectus|H. erectus]]'' (varian Afrika kadang-kadang disebut ''[[Homo ergaster|H. ergaster]]'') berevolusi 2 juta tahun yang lalu dan merupakan spesies [[manusia purba]] pertama yang meninggalkan Afrika dan menyebar ke seluruh Eurasia.<ref>{{cite journal|display-authors=6|date=July 2018|title=Hominin occupation of the Chinese Loess Plateau since about 2.1 million years ago|journal=[[Nature (journal)|Nature]]|volume=559|issue=7715|pages=608–612|bibcode=2018Natur.559..608Z|doi=10.1038/s41586-018-0299-4|pmid=29995848|vauthors=Zhu Z, Dennell R, Huang W, Wu Y, Qiu S, Yang S, Rao Z, Hou Y, Xie J, Han J, Ouyang T|s2cid=49670311}}</ref> ''H. erectus'' juga merupakan yang pertama kali mengembangkan bentuk tubuh yang khas sebagaimana manusia. ''Homo sapiens'' muncul di Afrika sekitar 300.000 tahun yang lalu berevolusi dari spesies yang umumnya dikenali sebagai ''[[Homo heidelbergensis|H. heidelbergensis]]'' atau ''[[Homo rhodesiensis|H. rhodesiensis]]'', yakni keturunan ''H. erectus'' yang masih ada di Afrika.<ref>{{cite journal|display-authors=6|date=June 2017|title=New fossils from Jebel Irhoud, Morocco and the pan-African origin of Homo sapiens|url=https://kar.kent.ac.uk/62267/1/Submission_288356_1_art_file_2637492_j96j1b.pdf|journal=[[Nature (journal)|Nature]]|volume=546|issue=7657|pages=289–292|bibcode=2017Natur.546..289H|doi=10.1038/nature22336|pmid=28593953|archive-url=https://web.archive.org/web/20200108234003/https://kar.kent.ac.uk/62267/1/Submission_288356_1_art_file_2637492_j96j1b.pdf|archive-date=8 January 2020|access-date=30 July 2022|vauthors=Hublin JJ, Ben-Ncer A, Bailey SE, Freidline SE, Neubauer S, Skinner MM, Bergmann I, Le Cabec A, Benazzi S, Harvati K, Gunz P|url-status=live}}</ref> ''H. sapiens'' bermigrasi keluar dari benua tersebut, secara bertahap menggantikan atau kawin silang dengan populasi manusia purba setempat.<ref>{{cite journal|author=<!--Staff writer(s); no by-line.-->|date=13 May 2005|title=Out of Africa Revisited|journal=[[Science (journal)|Science]]|type=This Week in ''Science''|volume=308|issue=5724|page=921|doi=10.1126/science.308.5724.921g|issn=0036-8075|s2cid=220100436}}</ref><ref>{{cite journal|author-link=Chris Stringer|date=June 2003|title=Human evolution: Out of Ethiopia|journal=[[Nature (journal)|Nature]]|volume=423|issue=6941|pages=692–3, 695|bibcode=2003Natur.423..692S|doi=10.1038/423692a|pmid=12802315|vauthors=Stringer C|s2cid=26693109}}</ref><ref>{{cite web|author-link=Donald Johanson|date=May 2001|title=Origins of Modern Humans: Multiregional or Out of Africa?|url=https://www.actionbioscience.org/evolution/johanson.html|website=[[actionbioscience]]|publisher=[[American Institute of Biological Sciences]]|location=Washington, DC|archive-url=https://web.archive.org/web/20210617010349/http://www.actionbioscience.org/evolution/johanson.html|archive-date=17 June 2021|access-date=23 November 2009|vauthors=Johanson D|url-status=dead}}</ref> Manusia mulai menunjukkan [[Modernitas perilaku|perilaku modern]] sekitar 160.000-70.000 tahun yang lalu,<ref name="Marean et al 2007">{{cite journal|last1=Marean|first1=Curtis|display-authors=etal|date=2007|title=Early human use of marine resources and pigment in South Africa during the Middle Pleistocene|url=http://doc.rero.ch/record/15550/files/PAL_E2962.pdf|journal=[[Nature (journal)|Nature]]|volume=449|issue=7164|pages=905–908|bibcode=2007Natur.449..905M|doi=10.1038/nature06204|pmid=17943129|s2cid=4387442}}</ref> dan mungkin lebih awal.<ref name="Brooks">{{Cite journal|year=2018|title=Long-distance stone transport and pigment use in the earliest Middle Stone Age|journal=[[Science (journal)|Science]]|volume=360|issue=6384|pages=90–94|bibcode=2018Sci...360...90B|doi=10.1126/science.aao2646|pmid=29545508|vauthors=Brooks AS, Yellen JE, Potts R, Behrensmeyer AK, Deino AL, Leslie DE, Ambrose SH, Ferguson JR, d'Errico F, Zipkin AM, Whittaker S, Post J, Veatch EG, Foecke K, Clark JB|doi-access=free}}</ref>
 
[[Asal-usul manusia modern dari Afrika|Migrasi "keluar dari Afrika"]] terjadi setidaknya dalam dua gelombang, gelombang pertama sekitar 130.000 hingga 100.000 tahun yang lalu, gelombang kedua (Penyebaran Selatan) sekitar 70.000 hingga 50.000 tahun yang lalu.<ref name="Posth">{{cite journal|display-authors=6|date=March 2016|title=Pleistocene Mitochondrial Genomes Suggest a Single Major Dispersal of Non-Africans and a Late Glacial Population Turnover in Europe|journal=[[Current Biology]]|volume=26|issue=6|pages=827–33|doi=10.1016/j.cub.2016.01.037|pmid=26853362|vauthors=Posth C, Renaud G, Mittnik A, Drucker DG, Rougier H, Cupillard C, Valentin F, Thevenet C, Furtwängler A, Wißing C, Francken M, Malina M, Bolus M, Lari M, Gigli E, Capecchi G, Crevecoeur I, Beauval C, Flas D, Germonpré M, van der Plicht J, Cottiaux R, Gély B, Ronchitelli A, Wehrberger K, Grigorescu D, Svoboda J, Semal P, Caramelli D, Bocherens H, Harvati K, Conard NJ, Haak W, Powell A, Krause J|hdl-access=free|s2cid=140098861|hdl=2440/114930}}</ref><ref>{{cite journal|display-authors=6|date=April 2015|title=A recent bottleneck of Y chromosome diversity coincides with a global change in culture|journal=[[Genome Research]]|volume=25|issue=4|pages=459–66|doi=10.1101/gr.186684.114|pmc=4381518|pmid=25770088|vauthors=Karmin M, Saag L, Vicente M, Wilson Sayres MA, Järve M, Talas UG, Rootsi S, Ilumäe AM, Mägi R, Mitt M, Pagani L, Puurand T, Faltyskova Z, Clemente F, Cardona A, Metspalu E, Sahakyan H, Yunusbayev B, Hudjashov G, DeGiorgio M, Loogväli EL, Eichstaedt C, Eelmets M, Chaubey G, Tambets K, Litvinov S, Mormina M, Xue Y, Ayub Q, Zoraqi G, Korneliussen TS, Akhatova F, Lachance J, Tishkoff S, Momynaliev K, Ricaut FX, Kusuma P, Razafindrazaka H, Pierron D, Cox MP, Sultana GN, Willerslev R, Muller C, Westaway M, Lambert D, Skaro V, Kovačevic L, Turdikulova S, Dalimova D, Khusainova R, Trofimova N, Akhmetova V, Khidiyatova I, Lichman DV, Isakova J, Pocheshkhova E, Sabitov Z, Barashkov NA, Nymadawa P, Mihailov E, Seng JW, Evseeva I, Migliano AB, Abdullah S, Andriadze G, Primorac D, Atramentova L, Utevska O, Yepiskoposyan L, Marjanovic D, Kushniarevich A, Behar DM, Gilissen C, Vissers L, Veltman JA, Balanovska E, Derenko M, Malyarchuk B, Metspalu A, Fedorova S, Eriksson A, Manica A, Mendez FL, Karafet TM, Veeramah KR, Bradman N, Hammer MF, Osipova LP, Balanovsky O, Khusnutdinova EK, Johnsen K, Remm M, Thomas MG, Tyler-Smith C, Underhill PA, Willerslev E, Nielsen R, Metspalu M, Villems R, Kivisild T}}</ref> H. sapiens kemudian menjajah semua benua dan pulau-pulau besar, tiba di [[Eurasia]] 125.000 tahun yang lalu,<ref>{{cite journal|date=January 2011|title=The southern route "out of Africa": evidence for an early expansion of modern humans into Arabia|url=https://www.sciencenews.org/view/generic/id/69197/title/Hints_of_earlier_human_exit_from_Africa|journal=[[Science (journal)|Science]]|volume=331|issue=6016|pages=453–6|bibcode=2011Sci...331..453A|doi=10.1126/science.1199113|pmid=21273486|archive-url=https://web.archive.org/web/20110427201317/https://www.sciencenews.org/view/generic/id/69197/title/Hints_of_earlier_human_exit_from_Africa|archive-date=27 April 2011|access-date=1 May 2011|vauthors=Armitage SJ, Jasim SA, Marks AE, Parker AG, Usik VI, Uerpmann HP|url-status=live|s2cid=20296624}}</ref><ref>{{cite web|date=27 January 2011|title=Humans 'left Africa much earlier'|url=https://www.bbc.co.uk/news/science-environment-12300228|work=[[BBC News]]|archive-url=https://web.archive.org/web/20120809051349/https://www.bbc.co.uk/news/science-environment-12300228|archive-date=9 August 2012|vauthors=Rincon P}}</ref> Australia sekitar 65.000 tahun yang lalu,<ref>{{cite journal|display-authors=6|date=July 2017|title=Human occupation of northern Australia by 65,000 years ago|journal=[[Nature (journal)|Nature]]|volume=547|issue=7663|pages=306–310|bibcode=2017Natur.547..306C|doi=10.1038/nature22968|pmid=28726833|vauthors=Clarkson C, Jacobs Z, Marwick B, Fullagar R, Wallis L, Smith M, Roberts RG, Hayes E, Lowe K, Carah X, Florin SA, McNeil J, Cox D, Arnold LJ, Hua Q, Huntley J, Brand HE, Manne T, Fairbairn A, Shulmeister J, Lyle L, Salinas M, Page M, Connell K, Park G, Norman K, Murphy T, Pardoe C|s2cid=205257212|hdl=2440/107043}}</ref> Amerika sekitar 15.000 tahun yang lalu, dan pulau-pulau terpencil seperti [[Hawaii]], [[Pulau Paskah]], [[Madagaskar]], dan [[Selandia Baru]] antara tahun 300 hingga 1280 M.<ref name="Lowe">{{cite web|year=2008|title=Polynesian settlement of New Zealand and the impacts of volcanism on early Maori society: an update|url=https://researchcommons.waikato.ac.nz/bitstream/10289/2690/1/Lowe%202008%20Polynesian%20settlement%20guidebook.pdf|publisher=[[University of Waikato]]|archive-url=https://web.archive.org/web/20100522032853/https://researchcommons.waikato.ac.nz/bitstream/10289/2690/1/Lowe%202008%20Polynesian%20settlement%20guidebook.pdf|archive-date=22 May 2010|access-date=29 April 2010|vauthors=Lowe DJ|url-status=live}}</ref><ref>{{cite journal|date=May 2012|title=Human migrations: Eastern odyssey|journal=[[Nature (journal)|Nature]]|volume=485|issue=7396|pages=24–6|bibcode=2012Natur.485...24A|doi=10.1038/485024a|pmid=22552074|vauthors=Appenzeller T|doi-access=free}}</ref>