Metabolisme: Perbedaan antara revisi

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=== Energi dari senyawa anorganik ===
{{further|Metabolisme mikrobial|Siklus nitrogen}}[[Litotrof|Kemolitotrof]] adalah jenis metabolisme yang ditemukan pada prokariota yang menggunakan energi yang didapatkan dari proses oksidasi [[senyawa anorganik]]. Organisme ini dapat menggunakan [[hidrogen]],<ref>{{Cite journal|last=Friedrich|first=B.|last2=Schwartz|first2=E.|date=1993-10-01|title=Molecular biology of hydrogen utilization in aerobic chemolithotrophs|url=https://www.annualreviews.org/doi/10.1146/annurev.mi.47.100193.002031|journal=Annual Review of Microbiology|volume=47|issue=1|pages=351–383|doi=10.1146/annurev.mi.47.100193.002031|issn=0066-4227|access-date=2020-07-02|archive-date=2022-07-19|archive-url=https://web.archive.org/web/20220719015853/https://www.annualreviews.org/doi/10.1146/annurev.mi.47.100193.002031|dead-url=yes}}</ref> senyawa [[belerang]] yang tereduksi ([[sulfida]], [[hidrogen sulfida]] dan [[tiosulfat]]),<ref>{{Cite book|url=http://www.sciencedirect.com/science/article/pii/S0065291108600181|title=Advances in Microbial Physiology|last=Friedrich|first=Cornelius G.|date=-01-01 1997|publisher=Academic Press|isbn=978-0-12-027739-1|editor-last=Poole|editor-first=R. K.|volume=39|location=|pages=235–289|language=en|doi=10.1016/s0065-2911(08)60018-1|issn=0065-2911|url-status=live|access-date=2020-07-02|archive-date=2015-09-24|archive-url=https://web.archive.org/web/20150924161219/http://www.sciencedirect.com/science/article/pii/S0065291108600181|dead-url=no}}</ref> [[Besi (II) oksida]]|besi (FeIIII) oksida]]<ref>{{Cite journal|last=Weber|first=Karrie A.|last2=Achenbach|first2=Laurie A.|last3=Coates|first3=John D.|date=01-10-2006|year=|title=Microorganisms pumping iron: anaerobic microbial iron oxidation and reduction|url=http://www.nature.com/articles/nrmicro1490|journal=Nature Reviews Microbiology|language=en|volume=4|issue=10|pages=752–764|doi=10.1038/nrmicro1490|issn=1740-1526|access-date=2020-07-02|archive-date=2021-03-08|archive-url=https://web.archive.org/web/20210308082034/https://www.nature.com/articles/nrmicro1490|dead-url=no}}</ref> atau [[amonia]]k<ref>{{Cite journal|last=Jetten|first=Mike S.M.|last2=Strous|first2=Marc|last3=van de Pas-Schoonen|first3=Katinka T.|last4=Schalk|first4=Jos|last5=van Dongen|first5=Udo G.J.M.|last6=van de Graaf|first6=Astrid A.|last7=Logemann|first7=Susanne|last8=Muyzer|first8=Gerard|last9=van Loosdrecht|first9=Mark C.M.|date=1998-12-01|year=|title=The anaerobic oxidation of ammonium|url=https://academic.oup.com/femsre/article-lookup/doi/10.1111/j.1574-6976.1998.tb00379.x|journal=FEMS Microbiology Reviews|language=en|volume=22|issue=5|pages=421–437|doi=10.1111/j.1574-6976.1998.tb00379.x|issn=1574-6976|access-date=2020-07-02|archive-date=2021-12-31|archive-url=https://web.archive.org/web/20211231003532/https://academic.oup.com/femsre/article-lookup/doi/10.1111/j.1574-6976.1998.tb00379.x|dead-url=no}}</ref> sebagai sumber energi dengan mengoksidasi senyawa tersebut dengan elektron aseptor, seperti oksigen atau [[nitrit]].<ref>{{Cite journal|last=Simon|first=Jörg|date=2002-08-01|year=|title=Enzymology and bioenergetics of respiratory nitrite ammonification|url=https://academic.oup.com/femsre/article-lookup/doi/10.1111/j.1574-6976.2002.tb00616.x|journal=FEMS Microbiology Reviews|language=en|volume=26|issue=3|pages=285–309|doi=10.1111/j.1574-6976.2002.tb00616.x|issn=1574-6976|access-date=2020-07-02|archive-date=2021-10-19|archive-url=https://web.archive.org/web/20211019061403/https://academic.oup.com/femsre/article-lookup/doi/10.1111/j.1574-6976.2002.tb00616.x|dead-url=no}}</ref> Proses mikrobial ini penting bagi [[daur biogeokimia]] , seperti [[asetogenesis]], [[nitrifikasi]] dan [[denitrifikasi]] sekaligus berfungsi penting untuk [[kesuburan tanah]] <ref>{{Cite journal|last=Barea|first=José-Miguel|last2=Pozo|first2=María José|last3=Azcón|first3=Rosario|last4=Azcón-Aguilar|first4=Concepción|date=2005-07-01|title=Microbial co-operation in the rhizosphere|url=http://academic.oup.com/jxb/article/56/417/1761/484466/Microbial-cooperation-in-the-rhizosphere|journal=Journal of Experimental Botany|language=en|volume=56|issue=417|pages=1761–1778|doi=10.1093/jxb/eri197|issn=1460-2431|access-date=2020-07-02|archive-date=2021-12-31|archive-url=https://web.archive.org/web/20211231003434/https://academic.oup.com/jxb/article/56/417/1761/484466/Microbial-cooperation-in-the-rhizosphere|dead-url=no}}</ref><ref>{{Cite journal|last=Conrad|first=R|date=1996-12-01|year=|title=Soil microorganisms as controllers of atmospheric trace gases (H2, CO, CH4, OCS, N2O, and NO).|url=https://mmbr.asm.org/content/60/4/609|journal=Microbiological reviews|language=en|volume=60|issue=4|pages=609–640|doi=10.1128/MMBR.60.4.609-640.1996|issn=0146-0749|access-date=2020-07-02|archive-date=2020-07-02|archive-url=https://web.archive.org/web/20200702162457/https://mmbr.asm.org/content/60/4/609|dead-url=no}}</ref>
 
=== Energi dari cahaya ===
{{further|Fototrof|Fotofosforilasi|Kloroplas}}Energi yang berasal dari sinar matahari ditangkap oleh [[tumbuhan]], [[sianobakteri]], [[bakteri ungu]],[[chlorobi]] dan beberapa jenis [[protista]]. Proses ini dipasangkan dengan konversi karbondioksida menjadi senyawa organik sebagai bagian proses fotosintesis. Penangkapan energi dan fiksasi karbon bisa beroperasi secara terpisah pada prokariota , sedangkan bakteri ungu fdan chlorobi bisa menggunakan sinar matahari sebagai sumber energi , ketika menukar reaksi di antara reaksi fiksasi karbon atau fermentasi senyawa organik.<ref>{{Cite journal|last=van der Meer|first=Marcel T. J.|last2=Schouten|first2=Stefan|last3=Bateson|first3=Mary M.|last4=Nübel|first4=Ulrich|last5=Wieland|first5=Andrea|last6=Kühl|first6=Michael|last7=de Leeuw|first7=Jan W.|last8=Sinninghe Damsté|first8=Jaap S.|last9=Ward|first9=David M.|date=2005-07|title=Diel Variations in Carbon Metabolism by Green Nonsulfur-Like Bacteria in Alkaline Siliceous Hot Spring Microbial Mats from Yellowstone National Park|url=https://aem.asm.org/content/71/7/3978|journal=Applied and Environmental Microbiology|language=en|volume=71|issue=7|pages=3978–3986|doi=10.1128/AEM.71.7.3978-3986.2005|issn=0099-2240|pmc=PMC1168979|pmid=16000812|access-date=2020-07-03|archive-date=2020-07-04|archive-url=https://web.archive.org/web/20200704064331/https://aem.asm.org/content/71/7/3978|dead-url=yes}}</ref><ref>{{Cite journal|last=Tichi|first=Mary A.|last2=Tabita|first2=F. Robert|date=2001-11-01|title=Interactive Control of Rhodobactercapsulatus Redox-Balancing Systems during Phototrophic Metabolism|url=https://jb.asm.org/content/183/21/6344|journal=Journal of Bacteriology|language=en|volume=183|issue=21|pages=6344–6354|doi=10.1128/JB.183.21.6344-6354.2001|issn=1098-5530|pmc=PMC100130|pmid=11591679|access-date=2020-07-03|archive-date=2021-03-08|archive-url=https://web.archive.org/web/20210308071829/https://jb.asm.org/content/183/21/6344|dead-url=yes}}</ref>
 
Pada banyak organisme, penangkapan energi sinar matahari memilik prinsip yang sama dengan fosforilasi oksidatif yang melibatkan penyimpanan energi sebagai gradien konsentrasi proton. Gaya gerak proton ini lah yang akan menggerakan sintesis ATP<ref>{{Cite journal|last=Alberts|first=Bruce|last2=Johnson|first2=Alexander|last3=Lewis|first3=Julian|last4=Raff|first4=Martin|last5=Roberts|first5=Keith|last6=Walter|first6=Peter|date=2002|title=Energy Conversion: Mitochondria and Chloroplasts|url=https://www.ncbi.nlm.nih.gov/books/NBK21063/|journal=Molecular Biology of the Cell. 4th edition|language=en|access-date=2020-07-03|archive-date=2020-12-15|archive-url=https://web.archive.org/web/20201215131416/https://www.ncbi.nlm.nih.gov/books/NBK21063/|dead-url=no}}</ref>.Elektron yang dibutuhkan untuk menggerakan rantai transpor elektron berasal dari protein yang mengumpulkan cahaya yang disebut [[pusat reaksi fotosintetikfotosintesis]]. Kelompok pusat reaksi ini dibagi menjadi dua tipe tergantung sifat [[Pigmen fotosintesis|pigmen fotosintetikfotosintesisnya]]nya di mana bakteri hanya memiliki satu tipe, sedangkan tumbuhan dan sianobakteri punya dua<ref>{{Cite journal|last=Allen|first=J. P.|last2=Williams|first2=J. C.|date=08-12-1998|year=|title=Photosynthetic reaction centers|url=https://febs.onlinelibrary.wiley.com/doi/abs/10.1016/S0014-5793%2898%2901245-9|journal=FEBS Letters|language=en|volume=438|issue=1-2|pages=5–9|doi=10.1016/S0014-5793(98)01245-9|issn=1873-3468|access-date=2020-07-03|archive-date=2022-04-11|archive-url=https://web.archive.org/web/20220411023011/https://febs.onlinelibrary.wiley.com/doi/abs/10.1016/S0014-5793%2898%2901245-9|dead-url=yes}}</ref>
 
Pada tumbuhan, alga, dan sianobakteri, [[fotosistem II]] menggunakan energi cahaya untuk melepaskan elektron dari molekul air dan melepaskan oksigen sebagai zat sisa. Lalu, elektron masuk ke dalam [[Kompleks sitokron b6F|kompleks sitokrom b6f]] yang menggunakan energi tersebut untuk memompa proten melewati membran [[tilakoid]] yang ada di dalam membran kloroplas. .<ref>{{Cite journal|last=Nelson|first=Nathan|last2=Ben-Shem|first2=Adam|date=2004-12-01|year=|title=The complex architecture of oxygenic photosynthesis|url=http://www.nature.com/articles/nrm1525|journal=Nature Reviews Molecular Cell Biology|language=en|volume=5|issue=12|pages=971–982|doi=10.1038/nrm1525|issn=1471-0072|access-date=2020-07-03|archive-date=2021-03-08|archive-url=https://web.archive.org/web/20210308134447/https://www.nature.com/articles/nrm1525|dead-url=no}}</ref> Proton ini kembali melalui membran untuk menggerakan ATP sintase seperti sebelumnya.Elektron masuk melalui [[fotosistem I]] dan bisa juga digunakan untuk mereduksi koenzim NADP<sup>+</sup><ref>{{Cite journal|last=Munekage|first=Yuri|last2=Hashimoto|first2=Mihoko|last3=Miyake|first3=Chikahiro|last4=Tomizawa|first4=Ken-Ichi|last5=Endo|first5=Tsuyoshi|last6=Tasaka|first6=Masao|last7=Shikanai|first7=Toshiharu|date=2004-06|title=Cyclic electron flow around photosystem I is essential for photosynthesis|url=http://www.nature.com/articles/nature02598|journal=Nature|language=en|volume=429|issue=6991|pages=579–582|doi=10.1038/nature02598|issn=0028-0836|access-date=2020-07-03|archive-date=2022-08-03|archive-url=https://web.archive.org/web/20220803101546/https://www.nature.com/articles/nature02598|dead-url=no}}</ref> Koenzim ini dapat digunakan di dalam [[siklus Calvin]] atau didaur ulang untuk menghasilkan ATP selanjutnya.